Los peces asociados a arrecifes del Parque Nacional Revillagigedo (Pacífico mexicano): un inventario curado, comentado e ilustrado

Autores/as

DOI:

https://doi.org/10.22201/ib.20078706e.2026.97.5757

Palabras clave:

Pacífico oriental tropical, Isla oceánica aislada, Arrecifes rocosos, Elasmobranquios, Peces óseos

Resumen

Accurate inventories of species living at isolated islands are essential for understanding the biogeographic and functional composition of insular faunas, habitat area vs. faunal-size relations and levels of endemism, as well as for purposes of conservation and management. Constructing such inventories involves new field research and rigorous winnowing of existing records that have errors in specimen identification and metadata or are unsubstantiable due to a lack of reliable source information. This paper provides a curated, photographically illustrated inventory of reef-associated fishes found at the 4 islands in the Revillagigedo National Park (RNP), 400 km southwest of Baja California. We assessed the quality of data about records relating to 364 species and added new data (specimens of 132 species and images of 182 species) that confirm species occurrences at each island in the park. The resultant inventory includes 234 accepted species occurrences, with 121 unaccepted occurrences (33.2% of the assessed species). Among the confirmed species, 166-189 (70.9-80.8%) evidently are park residents. New information indicates that the RNP fauna includes 23 named and probable species of endemics, representing 9.8% of the accepted species and 12.2-13.9% of the residents, highlighting the uniqueness of the RNP and its importance for conservation. 

Citas

Abesamis, R., Clements, K. D., Choat, J. H., McIlwain, J., Myers, R., Rocha, L. A. et al. (2012). Acanthurus xanthopterus. The IUCN Red List of Threatened Species 2012: e.T177989A1512937. Retrieved on August 20, 2024 from: http://dx.doi.org/10.2305/IUCN.UK.2012.RLTS.T177989A1512937.en

Aburto-Oropeza, O., Ballesteros, E., Ezcurra, E., Friedlander, A., Henning, B., Hoyos, M. et al. (2017). Archipiélago de Revillagigedo: biodiversidad, amenazas y necesidades de conservación. San Diego, California: UC San Diego, Aburto Lab.

Aburto-Oropeza, O., & Hull, P. M. (2008). A probable spawning aggregation of the leather bass Dermatolepis dermatolepis (Boulenger) in the Revillagigedo Archipelago, México. Journal of Fish Biology, 73, 288–295. https://doi.org/10.1111/j.1095-8649.2008.01909.x

Acevedo-Álvarez, E. A., Ruiz-Campos, G., & Domínguez-Domínguez, O. (2021). Population-level morphological variation of Anisotremus interruptus (Gill, 1862) (Perciformes: Haemulidae) in the Tropical Eastern Pacific, with the description of two new species. Zootaxa, 4975, 141–158. https://doi.org/10.11646/zootaxa.4975.1.5

Allen, G. R. (1981). Butterfly and angelfishes of the world, Vol. 2 Second Edition. Sydney, Australia: Mergus.

Allen, G. R., & Erdmann, M. V. (2024). Reef fishes of the East Indies, Second Edition, Volumes I–III. Perth, Australia: Tropical Reef Research.

Allen, G. R., & Randall, J. E. (1977). Review of the sharpnose pufferfishes (subfamily Canthigasterinae) of the Indo-Pacific. Records of the Australian Museum, 30, 475–511. https://doi.org/10.3853/j.0067-1975.30.1977.192

Allen, G. R., & Robertson, D. R. (1992a). Deux nouvelles especes de Girelles (Labridae: Halichoeres) du Pacifique oriental tropical. Revue française d’Aquariologie, 19, 47–52.

Allen, G. R., & Robertson, D. R. (1992b). Serranus socorroensis, a new species of serranid fish from the tropical eastern Pacific Ocean. Revue française d’Aquariologie, 19, 37–40.

Allen, G. R., & Robertson, D. R. (1992c). Three new species of Triplefins (Pisces: Tripterygiidae) from Malpelo and Socorro Islands, in the tropical eastern Pacific. Revue française d’Aquariologie, 19, 53–56.

Allen, G. R., & Robertson, D. R. (1997). An annotated checklist of the fishes of Clipperton Atoll, tropical eastern Pacific. Revista de Biologia Tropical, 45, 813–843

Allen, G. R., & Robertson, D. R. (1999). Epinephelus clippertonensis, a new species of serranid fish from the tropical eastern Pacific. Revue française d'Aquariologie Herpétologie, 26, 11–15.

Allen, G. R., & Woods, L. P. (1980). A review of the damselfish genus Stegastes from the eastern Pacific with the description of a new species. Records of the Western Australian Museum, 8, 171–198.

Anderson, A. B., Pinheiro, H. T., Batista, M. B., Francini-Filho, R. B., Gomes, L. E. O., Bernardino, A. F. et al. (2023). Biogeographic patterns of marine fishes associated with rhodolith beds in the Southwestern Atlantic reveal an ecotone of biodiversity. Biodiversity and Conservation, 32, 821–837. https://doi.org/10.1007/s10531-022-02528-0

Auster, P. J., Sánchez-Jiménez, A., Rodríguez-Arrieta, J. A., Quesada, A. J., Pérez, C., Naranjo-Elizondo, B. et al. (2016). Facilitative behavioral interactions between deepwater piscivores at Isla del Coco National Park and Las Gemelas Seamount, Costa Rica. Revista de Biología Tropical, 64, 187–196. https://doi.org/10.15517/rbt.v64i1.23425

Ayala-Bocos, A., Hoyos-Padilla, M., García-Benito, D., & Martínez-Castillo, V. (2015). New record of the frogfish Fowlerichthys avalonis (Actinopterygii, Antennariidae) at the oceanic Revillagigedo Archipelago, west Mexico. Marine Biodiversity Records, 8, 1–4. https://doi.org/10.1017/S1755267215000275

Berry, F. H., & Baldwin, W. J. (1966). Triggerfishes (Balistidae) of the eastern Pacific. Proceedings of the California Academy of Sciences (Series 4), 34, 429–474.

Baldwin, C. C., Tornabene, L., & Robertson, D. R. (2018). Below the mesophotic. Scientific Reports, 8, 4920. https://doi.org/10.1038/s41598-018-23067-1

Bautista-Romero, J., Reyes-Bonilla, D. B., & Lluch-Cota, S. E. (1994). Aspectos generales sobre la fauna marina. In A. Ortega-Rubio, & A. Castellanos-Vera (Eds.), La Isla Socorro, Reserva de la Biosfera Archipiélago Revillagigedo, México Publ. Núm. 8 (pp. 247–276). La Paz, México: CIBNOR.

Beebe, W. (1937). The Templeton Crocker Expedition. II. Introduction, itinerary, list of stations, nets and dredges. Zoologica, 22, 33–46.

Becerril-García, E. E., Gutiérrez-Ortiz, M. A., Preciado-González, P. A., & Ayala-Bocos, A. (2019). Presence of Remora remora on Mobula birostris in Revillagigedo National Park, Mexico. Marine and Freshwater Research, 71, 414–417. https://doi.org/10.1071/MF19089

Becerril-García, E. E., Hoyos-Padilla, E. M., Henning, B., & Salinas-De León, P. (2020). Sharks, rays, and chimaeras of the Revillagigedo National Park: an update of new and confirmed records. Journal of Fish Biology, 97, 1228–1232. https://doi.org/10.1111/jfb.14457

Bender, M. G., Leprieur, F., Mouillot, D., Kulbicki, M., Parravicini, V., Pie, M. R. et al. (2017). Isolation drives taxonomic and functional nestedness in tropical reef fish faunas. Ecography, 40, 425–435. https://doi.org/10.1111/ecog.02293

Bender, M. G., Pie, M. R., Rezende, E. L., Mouillot, D., & Floeter, S. R. (2013). Biogeographic, historical and environmental influences on the taxonomic and functional structure of Atlantic reef fish assemblages. Global Ecology and Biogeography, 22, 1173–1182. https://doi.org/10.1111/geb.12099

Bernardi, G., Findley, L., & Rocha-Olivares, A. (2003). Vicariance and dispersal across Baja California in disjunct marine fish populations. Evolution, 57, 1599–1609. https://doi.org/10.1111/j.0014-3820.2003.tb00367.x

Bernal-Hernández, M. E., Beltrán-López, R. G., Robertson, D. R., Baldwin, C. C., Espinoza, E., Martínez-Gómez, J. E. et al. (2024). Cryptic diversity in Scorpaenodes xyris (Jordan & Gilbert 1882) (Scorpaeniformes: Scorpaenidae) throughout the Tropical Eastern Pacific. Journal of Molecular Evolution, 92, 842–860. https://doi.org/10.1007/s00239-024-10212-w

Borg, J. A., Dandria, D., Evans, J., Knittweis, L., & Schembri, P. J. (2023). A critical checklist of the marine fishes of Malta and surrounding waters. Diversity, 15, 225. https://doi.org/10.3390/d15020225

Böhlke, E. B., & Randall, J. E. (2000). A review of the moray eels (Anguilliformes: Muraenidae) of the Hawaiian Islands, with descriptions of two new species. Proceedings of the Academy of Natural Sciences of Philadelphia, 150, 203–278.

Briggs, J. C. (1951). A review of the clingfishes (Gobiesocidae) of the eastern Pacific with descriptions of new species. Proceedings of the California Zoological Club, 1, 57–108.

Briggs, J. C. (1955). A monograph of the clingfishes (Order Xenopterygii). Stanford Ichthyological Bulletin, 6, 1–224.

Briggs, J. C. (1974). Marine Zoogeography. New York: McGraw-Hill

Briggs, J. C., & Bowen, B. W. (2013). Marine shelf habitat: biogeography and evolution. Journal of Biogeography, 40, 1023–1035. https://doi.org/10.1111/jbi.12082

Britzke, R., Menezes, N. A., & Nirchio, M. (2019). Redescription of Mugil setosus Gilbert 1892 with comments on the occurrence of Mugil curema Valenciennes 1836 in the Pacific Ocean (Teleostei: Perciformes: Mugilidae). Zootaxa, 4671, 396–406. https://doi.org/10.11646/zootaxa.4671.3.5

Bussing, W. A. (1987). Los peces de la familia Labridae de la Costa Pacífica de Costa Rica. Revista de Biología Tropical, 33, 81–98.

Bussing, W. A. (1990). New species of gobiid fishes of the genera Lythrypnus, Elacatinus and Chriolepis from the eastern tropical Pacific. Revista de Biología Tropical, 38, 99–118.

Bussing, W. A. (1991). A new species of eastern Pacific moray eel (Pisces: Muraenidae). Revista de Biología Tropical, 39, 97–102.

Cambra, M., Lara-Lizardi, F., Peñaherrera-Palma, C., Hearn, A., Ketchum, J. T., Zarate, P. et al. (2021). A first assessment of the distribution and abundance of large pelagic species at Cocos Ridge seamounts (Eastern Tropical Pacific) using drifting pelagic baited remote cameras. Plos One, 16, e0244343. https://doi.org/10.1371/journal.pone.0244343

Carvalho-Filho, A., Oliveira, C., Maximiano, L., Tavera, J., Acero, A. P., & Marceniuk, A. P. (2022). Review of the Pareques acuminatus (Bloch & Schneider, 1801) species complex, with revalidation of Pareques lineatus (Cuvier, 1830) from the Western Atlantic (Percomorphacea: Sciaenidae). Zootaxa, 5195, 401–418. https://doi.org/10.11646/zootaxa.5195.5.1

Castañeda-Beltrán, E. (1988). Prospección de la fauna ictiológica de isla Clarión, México. Biología Marina, Secretaría de Marina, México, 4, 97–136.

Castro-Aguirre, J. L., & Balart, E. F. (2002). La ictiofauna de las Islas Revillagigedo y sus relaciones zoogeográficas, con comentarios acerca de su origen y evolución. In M. L. Lozano-Vilano (Ed.), Libro jubilar en Honor al Dr. Salvador Contreras Balderas (pp. 153–170). Monterrey: Facultad de Ciencias Biológicas, Universidad Autónoma de Nuevo León.

Catanese, G., Manchado, M., & Infante, C. (2010). Evolutionary relatedness of mackerels of the genus Scomber based on complete mitochondrial genomes: strong support to the recognition of Atlantic Scomber colias and Pacific Scomber japonicus as distinct species. Gene, 452, 35–43. https://doi.org/10.1016/j.gene.2009.12.004

Chan, G. L. (1974). Report of biological observations of the Revillagigedo expedition. NAUI BioMarine Exploration Seminar, College of Maine, United States; unpublished, unreviewed report. https://www.biodiversitylibrary.org/bibliography/155008

Chao, N. L. (1995). Sciaenidae. In W. Fischer, F. Krupp, W. Schneider, C. Sommer, K. E. Carpenter, & V. H. Niem (Eds.), Guía FAO para la identificación para los fines de la pesca. Pacífico centro-oriental. Volumen III. Vertebrados - Parte 2 (pp. 1201–1813). Organización de las Naciones Unidas para la Agricultura y la Alimentación.

Chávez-Comparán, J. C., Patiño-Barragán, M., Calderón-Riveroll, G., Lezama-Cervantes, C., Lara-Chávez, B., Ibarra-Castillas, M. et al. (2010). Lista de peces generada por censos visuales submarinos en la isla Socorro Colima, México. Revista Cubana de Investigaciones Pesqueras, 27, 72–78.

Clements, K. D., Choat, J. H., Abesamis, R., McIlwain, J., Myers, R., Nanola, C. et al. (2012). Ctenochaetus marginatus. The IUCN Red List of Threatened Species 2012: e.T177964A1504292. https://dx.doi.org/10.2305/IUCN.UK.2012.RLTS.T177964A1504292.en

Collette, B. B., & Banford, H. M. (2001). Status of the eastern Pacific agujon needlefish Tylosurus pacificus (Steindachner, 1876) (Beloniformes: Belonidae). Revista de Biología Tropical, 49 (Suppl. 1), 51–57.

Collette, B., Polanco-Fernández, A., & Aiken, K. A. (2015). Ablennes hians. The IUCN Red List of Threatened Species 2015: e.T13486514A15603320. https://dx.doi.org/10.2305/IUCN.UK.2015-4.RLTS.T13486514A15603320.en

Collette, B. B., & Graves, J. (2019). Tunas and billfishes of the World. Johns Hopkins University Press, Baltimore, Maryland.

Collette, B. B., Nakatsuka, S., & Suzuki, J. (2023). Scomber japonicus. The IUCN Red List of Threatened Species 2023: https://dx.doi.org/10.2305/IUCN.UK.2023-1.RLTS.T170306A170083106.en

Conanp (Comisión Nacional de Áreas Naturales Protegidas). (2019). Programa de manejo Parque Nacional Revillagigedo. Mexico: Conanp.

Cord, I., Floeter, S. R., Araújo, G. S., Quimbayo, J. P., Robertson, D. R., Victor, B. C. et al. (2024). Atlantic reef fishes: distributions and life-history traits (Version 1) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.13655554

Dawson, C. E. (1975). Studies on eastern Pacific sand stargazers (Pisces: Dactyloscopidae). 2. Genus Dactyloscopus, with descriptions of new species and subspecies. Science Bulletin, Natural History Museum of Los Angeles County, 22, 1–61

Dawson, C. E. (1981). Review of the Indo-Pacific pipefish genus Doryrhamphus Kaup (Pisces: Syngnathidae), with descriptions of a new species and a new subspecies. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology, 44, 1–27

Dawson, C. E. (1985). Indo-Pacific pipefishes (Red Sea to the Americas). Gulf Coast Research Laboratory, Ocean Springs, Mississippi.

Del Moral-Flores, L. F., Negrete, J. M. G., & Camacho, A. F. G. (2016). Peces del archipiélago de las islas Revillagigedo: una actualización sistemática y biogeográfica. Biocyt: Biología, Ciencia y Tecnología, 9, 596–619.

Díaz-Licona, C. A., Schubert, N., González-Gamboa, V., Tuya, F., Azofeifa-Solano, J. C., & Fernández-García, C. (2025). Rhodolith beds in the Eastern Tropical Pacific: Habitat structure and associated biodiversity. Aquatic Botany, 201, 103914. https://doi.org/10.1016/j.aquabot.2025.103914

Dubuc, A., Quimbayo, J. P., Alvarado, J. J., Araya-Arce, T., Arriaga, A., Ayala-Bocos, A. et al. (2023). Patterns of reef fish taxonomic and functional diversity in the Eastern Tropical Pacific. Ecography, 2023, e06536. https://doi.org/10.1111/ecog.06536

Durand, J. D., & Borsa, P. (2015). Mitochondrial phylogeny of grey mullets (Acanthopterygii: Mugilidae) suggests high proportion of cryptic species. Comptes Rendus Biologies, 338, 266–277

Estape, A. M. (2023). Documenting reef-fish diversity in the Revillagigedo Archipelago, Pacific Mexico, November 2022. Zenodo. https://zenodo.org/records/8146184

Ferrari, D. S., Floeter, S. R., Leprieur, F., & Quimbayo, J. P. (2023). A trait-based approach to marine island biogeography. Journal of Biogeography, 50, 528–538. https://doi.org/10.1111/jbi.14549

Findley, L. T. (1983). A revision of the eastern Pacific species of the gobiid genus Chriolepis (Teleostei: Gobiidae) (Mexico, Latin America) (Ph.D. Dissertation). University of Arizona, Tucson AZ.

Fitch, J. E. (1956). Pacific mackerel. California Cooperative Oceanic Fisheries Investigations Progress Report, 1956, 29–32.

Fitch, J. E., & Crooke, S. J. (1984). Revision of eastern Pacific catalufas (Pisces: Priacanthidae) with description of a new genus and discussion of the fossil record. Proceedings of the California Academy of Sciences, 43, 301–315.

Fontes, J. T., Katoh, K., Pires, R., Soares, P., & Costa, F. O. (2024). Benchmarking the discrimination power of commonly used markers and amplicons in marine fish (e)DNA (meta)barcoding. Metabarcoding and Metagenomics, 8, 295–320. https://doi.org/10.3897/mbmg.8.128646

Fourriére, M., Alvarado, J. J., Ayala-Bocos, A., & Cortes, J. (2017). Updated checklist and analysis of completeness of the marine fish fauna of Isla del Coco, Pacific of Costa Rica. Marine Biodiversity, 47, 813–821. https://doi.org/10.1007/s12526-016-0501-6

Fourriére, M., Reyes-Bonilla, H., Ayala-Bocos, A., Ketchum, J. A., & Chávez-Comparan, J. C. (2016). Checklist and analysis of completeness of the reef fish fauna of the Revillagigedo Archipelago, Mexico. Zootaxa, 4150, 436–466. http://doi.org/10.11646/zootaxa.4150.4.4

Fourriére, M., Reyes-Bonilla, H., Rodríguez-Zaragoza, F. A., & Crane, N. (2014). Fishes of Clipperton Atoll, Eastern Pacific: checklist, endemism, and analysis of completeness of the inventory. Pacific Science, 68, 375–395. https://doi.org/10.2984/68.3.7

Fricke, R., Eschmeyer, W. N., & Van der Laan, R. (Eds.). (2026). Eschmeyer’s Catalog of fishes: genera, species, references. Electronic version last accessed on 1 March, 2026. http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp

Fricke, R., Reséndiz-López, M. A. & Oseguera-Rodríguez, A. S. (2024). Fishes and lampreys of Mexico. An annotated checklist. Mexico City: Conabio.

Fritzsche, R. A. (1980). Revision of the eastern Pacific Syngnathidae (Pisces: Syngnathiformes), including both recent and fossil forms. Proceedings of the California Academy of Sciences, 42, 181–227.

GBIF: Global Biodiversity Information Facility. https://www.gbif.org; last accessed on August 13, 2025.

Giddens, J., Goodell, W., Friedlander, A., Salinas-de-León, P., Shepard, C., Henning, B. et al. (2019). Patterns in bathyal demersal biodiversity and community composition around archipelagos in the Tropical Eastern Pacific. Frontiers in Marine Science, 6, 1–15. https://doi.org/10.3389/fmars.2019.00388

Gilbert, C. H. (1890). A preliminary report on the fishes collected by the steamer Albatross on the Pacific coast of North America during the year 1889, with descriptions of twelve new genera and ninety-two new species. Proceedings of the United States National Museum, 13, 49–126.

Gilbert, C. H. (1892). Descriptions of thirty-four new species of fishes collected in 1888 and 1889, principally among the Santa Barbara Islands and in the Gulf of California. In Scientific results of explorations by the U. S. Fish Commission Steamer Albatross (pp. 539–566). Proceedings of the United States National Museum, 14(No. 880).

Ginsburg, I. (1947). American species and subspecies of Bathygobius, with a demonstration of a suggested modified system of nomenclature. Journal of the Washington Academy of Sciences, 37, 275–284.

Gotshall, D. W. (1998). Sea of Cortez marine animals: a guide to the common fishes and invertebrates – Baja California to Panama. Monterey, CA: Sea Challengers.

Greenfield, D. W. (1965). Systematics and zoogeography of Myripristis in the eastern tropical Pacific. California Fish and Game, 51, 229–247.

Grove, J. S. & Lavenberg, R. J. (1997). The fishes of the Galápagos Islands. Stanford, CA: Stanford University Press.

Guimarães, R. Z. P. (1999). Revision, phylogeny and comments on biogeography of soapfishes of the genus Rypticus (Teleostei: Serranidae). Bulletin of Marine Science, 65, 337–379.

Hashimoto, S., & Motomura, H. (2025). First Japanese records of Priacanthus alalaua (Perciformes: Priacanthidae) from the Ryukyu Islands.Biogeography, 27, 47-51.

https://doi.org/10.11358/biogeo.27.47

Hastings, P. A., & Robertson, D. R. (1999). Acanthemblemaria atrata and Acanthemblemaria mangognatha, new species of eastern Pacific barnacle blennies (Chaenopsidae) from Isla del Coco, Costa Rica, and Islas Revillagigedo, Mexico, and their relationships with other barnacle blennies. Revue française d'Aquariologie Herpétologie, 25, 107–118.

Hastings, P. A., & Springer, V. G. (2009). Recognizing diversity in blennioid fish nomenclature (Teleostei: Blennioidei). Zootaxa, 2120, 3–14. https://doi.org/10.11646/zootaxa.2120.1.2

Heller, E., & Snodgrass, R. E. (1903). Papers from the Hopkins Stanford Galápagos expedition, 1898–1899. XV. New fishes. Proceedings of the Washington Academy of Sciences, 5, 189–229.

Herald, E. S. (1940). A key to the pipefishes of the Pacific American coasts with descriptions of new genera and species. Report Allan Hancock Pacific Expedition, 1932–1938, 9, 51–64.

Hobbs, J. P. A., Jones, G. P., & Munday, P. L. (2011). Extinction risk in endemic marine fishes. Conservation Biology, 25, 1053–1055. https://doi.org/10.1111/j.1523-1739.2011.01698.x

Hobbs, J. P. A., Jones, G. P., Munday, P. L., Connolly, S. T., & Srinivasan, M. (2012). Biogeography and the structure of coral reef fish communities on isolated islands. Journal of Biogeography, 39, 130–139. https://doi.org/10.1111/j.1365-2699.2011.02576.x

Hobbs, J. P. A., Newman, S. J., Mitsopoulos, G. E. A., Travers, M. J., Skepper, C. L., Gilligan, J. J. et al. (2014). Checklist and new records of Christmas Island fishes: the influence of isolation, biogeography and habitat availability on species abundance and community composition. Raffles Bulletin of Zoology (Suppl. 30), 184–202.

Hollarsmith, J. A., Winquist, R. O. T., Velasco-Lozano, M., DuBois, K., Reyes-Bonilla, H., Neumann, K. C. et al. (2020). Habitats and fish communities at mesophotic depths in the Mexican Pacific. Journal of Biogeography, 47, 1552–1563. https://doi.org/10.1111/jbi.13842

Hubbs, C. (1953). Revision of the eastern Pacific fishes of the clinid genus Labrisomus. Zoologica, Scientific Contributions of the New York Zoological Society, 38, 113–136.

Hunter, J. R., & Mitchell, C. T. (1966). Association of fishes with flotsam in offshore waters of Central America. US Fishery Bulletin, 66, 13–29.

Hunter, J. R., & Mitchell, C. T. (1968). Field experiments on the attraction of pelagic fish to floating objects. Journal du Conseil / Conseil Permanent International pour l'Exploration de la Mer, 31, 427–434.

iNaturalist. https://www.inaturalist.org; accessed many times between August, 2025 and March, 2026.

IOC-UNESCO. (2024). State of the Ocean Report. Paris: IOC-UNESCO. (IOC Technical Series, 190). https://doi.org/10.25607/4wbg-d349

Jordan, D. S., & Evermann, B. W. (1903). Descriptions of new genera and species of fishes from the Hawaiian Islands. Bulletin of the U. S. Fish Commission, 22, 161–208.

Jordan, D. S., & Gilbert, C. H. (1882). Notes on a collection of fishes made by Lieut. Henry E. Nichols, U. S. N., on the west coast of Mexico, with descriptions of new species. Proceedings of the United States National Museum, 4, 225–233.

Jordan, D. S., & McGregor, R. C. (1899). List of fishes collected at the Revillagigedo Archipelago and neighboring islands. United States Commission of Fish and Fisheries, Report of the Commissioner, 24, 271–284.

Klimley, A. P., Ketchum, J. T., Lara-Lizardi, F., Papastamatiou, Y. P., & Hoyos-Padilla, E. M. (2022). Evidence for spatial and temporal resource partitioning of sharks at Roca Partida, an isolated pinnacle in the eastern Pacific. Environmental Biology of Fishes, 105, 1963–1974. https://doi.org/10.1007/s10641-021-01206-4

Knudsen, S. W., & Clements, K. D. (2013). Revision of the family Kyphosidae (Teleostei: Perciformes). Zootaxa, 3751, 1–101. https://doi.org/10.11646/zootaxa.3751.1.1

Knudsen, S. W., & Clements, K. D. (2016). World-wide species distributions in the family Kyphosidae (Teleostei: Perciformes). Molecular Phylogenetics and Evolution, 101, 252–266. http://dx.doi.org/10.1016/j.ympev.2016.04.037

Konow, N., & Ferry-Graham, L. (2013). Functional morphology of butterflyfishes. In M. S. Pratchett, M. L. Berumen, & B. G. Kapoor (Eds.), The Biology of butterflyfishes (pp. 19–47). Boca Raton, Florida: CRC Press.

Krejsa, R. J. (1960). The eastern, tropical Pacific fishes of the genus Blenniolus, including a new island endemic. Copeia, 1960, 322–336. https://doi.org/10.2307/1439760

Lara-Lizardi, F., Hoyos-Padilla, M., Ketchum, J. T., & Galván-Magaña, F. (2018). Range expansion of the whitenose shark, Nasolamia velox, and migratory movements to the oceanic Revillagigedo Archipelago (west Mexico). Journal of the Marine Biological Association of the United Kingdom, 98, 949–953. https://doi.org/10.1017/S0025315417000108

Lea, R. N., Fraser, T. H., Baldwin, C. C., & Craig, M. T. (2022). Five valid species of cardinalfishes of the genus Apogon (Apogonidae) in the eastern Pacific Ocean, with a redescription of A. atricaudus and notes on the distribution of A. atricaudus and A. atradorsatus. Ichthyology & Herpetology, 110, 106–114. https://doi.org/10.1643/i2020137

Lessios, H. A., & Robertson, D. R. (2006). Crossing the impassable: genetic connections in 20 reef fishes across the Eastern Pacific Barrier. Proceedings of the Royal Society B, 273, 2201–2208.

Love, M. S., Bizzarro, J. J., Cornthwaite, A. M., Frable, B. W., & Maslenikov, K. P. (2021). Checklist of marine and estuarine fishes from the Alaska–Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico. Zootaxa, 5053, 1–285. https://doi.org/10.11646/zootaxa.5053.1.1

Ludt, W. B., Bernal, M. A., Kenworthy, E., Salas, E., & Chakrabarty, P. (2019). Genomic, ecological, and morphological approaches to investigating species limits: A case study in modern taxonomy from Tropical Eastern Pacific surgeonfishes. Ecology and Evolution, 9, 4001–4012. https://doi.org/10.1002/ece3.5029

Martínez-Takeshita, N., Purcell, C. M., Chabot, C. L., Craig, M. T., Paterson, C. N., Hyde, J. R. et al. (2015). A tale of three tails: cryptic speciation in a globally distributed marine fish of the genus Seriola. Copeia, 103, 357–368. https://doi.org/10.1643/CI-124-224

Mason, B. M., Mesaglio, T., Heitmann, J. B., Chandler, M., Chowdhury, S., Gorta, S. B. Z. et al. (2025). iNaturalist accelerates biodiversity research. BioScience, 2025, biaf104. https://doi.org/10.1093/biosci/biaf104

Matsuura, K. (2022). Family Balistidae, Triggerfishes. In P. C. Heemstra, E. Heemstra, D. A. Ebert, W. Holleman, & J. E. Randall (Eds.), Coastal fishes of the western Indian Ocean, Volume 5 (pp. 414–429). Makhanda, South Africa: South African Institute for Aquatic Biodiversity.

McCarthy, L. V. (1979). Eastern Pacific Rypticus (Pisces: Grammistidae). Copeia, 1979, 393–400. https://doi.org/10.2307/1443213

McCosker, J. E., & Rosenblatt, R. H. (2010). The fishes of the Galápagos Archipelago: an update. Proceedings of the California Academy of Sciences, 61, 167–195.

McPhail, J. D. (1961). A review of the Tropical Eastern Pacific species of Pareques (Sciaenidae). Copeia, 1961, 27–32. https://doi.org/10.2307/1440167

Medina-Rosas, P., & Moreno-López, J. A. (2025). First records of the Indo-Pacific Platax teira (Forsskål, 1775), Longfin Batfish (Perciformes, Ephippidae), in the Tropical Eastern Pacific. Check List, 21, 427–432. https://doi.org/10.15560/21.2.427

Miller, P. J., & Stefanni, S. (2001). The eastern Pacific species of Bathygobius (Perciformes: Gobiidae). Revista de Biología Tropical, 49 (Suppl. 1), 141–156.

Mora, C., Francisco, V., & Zapata, F. A. (2001). Dispersal of juvenile and adult reef fishes associated with floating objects and their recruitment into Gorgona Island reefs, Colombia. Bulletin of Marine Science, 68, 557–561.

Mundy, B. C. (2005). Checklist of the fishes of the Hawaiian Islands. Bishop Museum Bulletin in Zoology, 6, 1–703.

Mundy, B. C., Richard, W., Demartini, E., Greene, B., Zgliczynski, B., Schroeder, R. E. et al. (2010). Inshore fishes of Howland Island, Baker Island, Jarvis Island, Palmyra Atoll, and Kingman Reef. Atoll Research Bulletin, 585, 1–132. https://doi.org/10.5479/si.00775630.585

Myers, R., & Pratchett, M. (2010). Forcipiger longirostris. The IUCN Red List of Threatened Species 2010: e.T165667A6085300. http://dx.doi.org/10.2305/IUCN.UK.2010-4.RLTS.T165667A6085300.en

OBIS (Ocean Biodiversity Information System), https://obis.org/; Accessed August, 2025.

Ochoa-López, E., Villavicencio-Garayzar, C. J., & Ruiz-Gaytán, A. (1997). First record of whitetip reef shark, Triaenodon obesus on the west coast of México (Revillagigedo Archipelago). Revista de Biologia Tropical, 44/45, 687–688.

Olivier, D., Loiseau, N., Petatán-Ramírez, D., Millán, O. T., Suárez Castillo, A. N., Torre, J. et al. (2018). Functional biogeography of the reef fishes of the islands of the Gulf of California: Integrating functional divergence into marine conservation. Global Ecology and Conservation, 16, e00506. https://doi.org/10.1016/j.gecco.2018.e00506

Palacios-Morales, G., Torres-Hernández, E., Campos-Medina, A., & Domínguez-Domínguez, O. (2014). Taxonomic and genetic corroboration of the presence of the bridled trigger fish Sufflamen fraenatum (Balistidae) for the coast of Mexico. Revista Mexicana de Biodiversidad, 85, 641–644. https://doi.org/10.7550/rmb.36883

Palacios-Salgado, D. S., Campos-Dávila, L., Granados-Amores, J., Cruz-Escalona, V. H., Peterson, M. S., Moreno-Sánchez, X. G. et al. (2019). Functional diversity in fish assemblages of the Tropical Eastern Pacific Ocean: A review of two decades of progress in the functional diversity approach. Hidrobiológica, 29, 17–40. https://doi.org/10.24275/uam/izt/dcbs/hidro/2019v29n1/Palacios

Petatán-Ramírez, D., Webster, P., Boye, L., Vázquez-Vera, L., & Reyes-Bonilla, H. (2025). First record of Platax teira (Forsskål, 1775), Longfin Batfish (Acanthuriformes, Ephippidae), in Cabo Pulmo National Park, Mexico. Check List, 21, 534–538. https://doi.org/10.15560/21.3.534

Pietsch, T. W., & Arnold, R. J. (2020). Frogfishes. Biodiversity, zoogeography, and behavioral ecology. Baltimore: Johns Hopkins University Press.

Pinheiro, H. T., Rocha, L. A., Macieira, R. M., Carvalho-Filho, A., Anderson, A. B., Bender, M. G. et al. (2018). South-western Atlantic reef fishes: zoogeographical patterns and ecological drivers reveal a secondary biodiversity centre in the Atlantic Ocean. Diversity and Distributions, 24, 951–965. https://doi.org/10.1111/ddi.12729

Pinheiro, H. T., Shepherd, B., Castillo, C., Abesamis, R. A., Copus, J. M., Pyle, R. L. et al. (2019). Deep reef fishes in the world’s epicenter of marine biodiversity. Coral Reefs, 38, 985–995. https://doi.org/10.1007/s00338-019-01825-5

Ramírez-Zúñiga, M. A., Molina-Alonso, A., Carmona-Ruiz, Y. A., León-Solórzano, E., Salgado-Castrejón, M., Serviere-Zaragoza, E. et al. (2025). Arrecifes rocosos. In Área marina del Parque Nacional Revillagigedo: pasado, presente y futuro de la investigación marina. Retrieved on July 25 from https://escholarship.org/uc/item/3wp552dt

Randall, J. E. (2005). Reef and shore fishes of the South Pacific. New Caledonia to Tahiti and the Pitcairn Islands. Honolulu: University of Hawaii Press.

Randall, J. E. (2007). Reef and shore fishes of the Hawaiian Islands. Honolulu: Sea Grant College Program, University of Hawai'i.

Ricker, K. E. (1959). Fishes collected from the Revillagigedo Islands during the 1954–1958 cruises of the “Marijean”. Institute of Fisheries, University of British Columbia, Vancouver, Canada Museum Contribution, 4, 1–10.

Robertson, D. R. (2024). Reef-associated Bony Fishes of the Tropical Eastern Pacific: a Database (Version 1). Zenodo. https://doi.org/10.5281/zenodo.13863109

Robertson, D. R., & Allen, G. R. (1996). Zoogeography of the shorefish fauna of Clipperton Atoll. Coral Reefs, 15, 121–131.

Robertson, D. R., & Allen, G. R. (2006). Shorefishes of the Tropical Eastern Pacific: an information system. CD-ROM. Smithsonian Tropical Research Institute, Balboa, Panama (superseded, no longer available).

Robertson, D. R., & Allen, G. R. (2015). Shorefishes of the Tropical Eastern Pacific: online information system. Version 2.0. Smithsonian Tropical Research Institute, Balboa, Panama (superseded, no longer available).

Robertson, D. R., & Allen, G. R. (2024). Shorefishes of the Tropical Eastern Pacific: online information system. Version 3.0. Smithsonian Tropical Research Institute, Balboa, Panama. www.stri.org/sftep

Robertson, D. R., & Cramer, K. (2009). Marine shore-fishes and biogeographic subdivisions of the Tropical Eastern Pacific. Marine Ecology Progress Series, 380, 1–17.

Robertson, D. R., Tornabene, L., Lardizabal, C. C., & Baldwin, C. C. (2022). Submersibles greatly enhance research on the diversity of deep-reef fishes in the Greater Caribbean. Frontiers in Marine Science, 8, 800250. https://doi.org/10.3389/fmars.2021.800250

Rosenblatt, R. H., Miller, E. C., & Hastings, P. A. (2013). Three new species of triplefin blennies of the genus Enneanectes (Teleostei, Tripterygiidae) from the tropical eastern Pacific with a key to Pacific species of Enneanectes. Zootaxa, 3636, 361–373.

Sabaj, M. H. (2023). Codes for Natural History Collections in Ichthyology and Herpetology (online supplement). Version 9.5 (10 Nov 2023). American Society of Ichthyologists and Herpetologists, Washington, DC. Retrieved from https://asih.org

Sandin, S. A., Vermeij, M. J., & Hurlbert, A. H. (2008). Island biogeography of Caribbean coral reef fish. Global Ecology and Biogeography, 17, 770–777.

Sánchez-García, C., Escobar-Sánchez, O., Valdez-Pineda, M. C., Ramírez-Pérez, J. S., Morán-Angulo, R. E., & Moreno-Sánchez, X. G. (2017). Selective predation by crevalle jack Caranx caninus on engraulid fishes in the SE Gulf of California, Mexico. Environmental Biology of Fishes, 100, 899–912. https://doi.org/10.1007/s10641-017-0615-0

Schmitt, W. L., & Schultz, L. P. (1940). List of the fishes taken on the Presidential Cruise of 1938. Smithsonian Miscellaneous Collections, 98, 1–10.

Schultz, L. P. (1946). A revision of the genera of mullets, fishes of the family Mugilidae, with descriptions of three new genera. Proceedings of the United States National Museum, 96, 377–395.

Scoles, D. R., Collette, B. B., & Graves, J. E. (1998). Global phylogeography of mackerels of the genus Scomber. Fishery Bulletin, 96, 823–842.

Snodgrass, R. E., & Heller, E. (1905). Papers from the Hopkins-Stanford Galápagos Expedition, 1898–1899. XVII. Shore fishes of the Revillagigedo, Clipperton, Cocos and Galápagos Islands. Proceedings of the Washington Academy of Science, 6, 333–427.

Springer, V. G. (1962). A review of the blenniid fishes of the genus Ophioblennius Gill. Copeia, 1962, 426–433.

Starnes, W. C. (1988). Revision, phylogeny and biogeographic comments on the circumtropical marine percoid fish family Priacanthidae. Bulletin of Marine Science, 43, 117–203.

Stevens, G. (2023). https://www.mantatrust.org/. Accessed 2023.

Stiller, J., Short, G., Hamilton, H., Saarman, N., Longo, S., Wainwright, P. et al. (2022). Phylogenomic analysis of Syngnathidae reveals novel relationships, origins of endemic diversity and variable diversification rates. BMC Biology, 20, 75. https://doi.org/10.1186/s12915-022-01271-w

Tanner, Z. L. (1889). Report upon the investigations of the U.S. Fish Commission steamer Albatross for the year ending June 20, 1889. Report of the U.S. Commission of Fish and Fisheries for 1888 (1892), XVI, 395–512.

Taylor, B. M., Prince, J., Mutz, S., Pardee, C., Wiley, J., Robertson, D. R. et al. (2024). A widespread, consistent, and perplexing biphasic pattern in log catch-at-age data from a widely-harvested family of tropical reef fishes. Fish and Fisheries, 25, 910-917. https://doi.org/10.1111/faf.12853

Teague, G. W. (1951). The sea-robins of America: a revision of the triglid fishes of the genus Prionotus. Comunicaciones Zoológicas del Museo de Historia Natural de Montevideo, 3, 1–59.

Thieme, P., Reisser, C., Bouvier, C., Rieuvilleneuve, F., Béarez, P., Coleman, R. R. et al. (2025). Historical biogeography of the Mugil cephalus species complex and its rapid global colonization. Molecular Phylogenetics and Evolution, 205, 108296. https://doi.org/10.1016/j.ympev.2025.108296

Torres-García, R., Ramírez-Ortiz, G., De León-Siller, E., Beas-Luna, R., Lorda, J., Malpica-Cruz, L. et al. (2025). Taxonomic and functional richness of fish in temperate and tropical reefs of the Mexican Pacific. Ciencias Marinas, 50, e3512. https://doi.org/10.7773/cm.y2025.3512

UNESCO (Organización de las Naciones Unidas para la Educación, la Ciencia y la Cultura). (n.d.). eDNA expedition to the Archipiélago de Revillagigedo. OBIS database. Retrieved on 3 March, 2025, from https://obis.org/dataset/c8705088-d0be-4a84-985b-c2cabcbc0879

Valencia-Méndez, O., Domínguez-Domínguez, O., López-Pérez, A., Martínez-Gómez, J. E., & Ayala-Bocos, A. (2018). Partial albinism in the Revillagigedo sea chub Kyphosus sectatrix (Perciformes: Kyphosidae) from Clarion Island, Mexico. Revista Mexicana de Biodiversidad, 89, 572–576. https://doi.org/10.22201/ib.20078706e.2018.2.2242

Vaudo, J. J., Plum, T. S., Logan, R. K., Wetherbee, B. M., & Mahmood, S. S. (2023). Diversity and spatiotemporal variation of an Eastern Tropical Pacific pelagic vertebrate community assessed with drifting pelagic BRUVS. Marine Biology, 170, 157. https://doi.org/10.1007/s00227-023-04308-y

Velasco-Lozano, M. F., Ramírez-Ortiz, G., Reyes-Bonilla, H., & Hollarsmith, J. A. (2020). Fish assemblages at mesophotic depths in the Pacific: a comparison between continental and oceanic islands of Mexico. Ciencias Marinas, 46, 321–342. https://doi.org/10.7773/cm.v46i4.3112

Victor, B. C., Frable, B. W., & Ludt, W. B. (2024). Halichoeres sanchezi n. sp., a new wrasse from the Revillagigedo Archipelago of Mexico, tropical eastern Pacific Ocean (Teleostei: Labridae). PeerJ, 12, e16828. https://doi.org/10.7717/peerj.16828

Victor, B. C., Grove, J. S., Long, D. J., Robertson, D. R., Keith, I., Bensted-Smith, W., & Salinas-de-León, P. (2024). List of Fishes of the Galápagos Archipelago, Ecuador (Version 2.0). Journal of the Ocean Science Foundation, 41, 54–111. https://doi.org/10.5281/zenodo.14057695

Victor, B. C., Wellington, G. M., Robertson, D. R., & Ruttenberg, B. (2001). Effects of the El Niño-Southern Oscillation event on the distribution of reef associated labrid fishes in the eastern Pacific Ocean. Bulletin of Marine Science, 69, 279–288.

Zenetos, A., Corsini-Foka, M., Crocetta, F., Gerovasileiou, V., Karachle, V., Simboura, M. et al. (2018). Deep cleaning of alien and cryptogenic species records in the Greek Seas (2018 update). Management of Biological Invasions, 9, 209–226. https://doi.org/10.3391/mbi.2018.9.3.04

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2026-08-20

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Robertson, D. R., Frable , B., Victor, B. C., Ludt, W., Gaither, M. R., Estape, C. J., Morgan-Estape, A., Pitassy, D. ., Ilves, K. L., Domínguez-Domínguez, O., Valencia-Méndez, O., Ayala-Bocos, A., Acevedo-Álvarez , E. A., Solís-Guzmán , M. G. ., Awhida-Robinson , A. K. ., Martínez-García , L. F. ., Ramos-Mayoral , A. G. ., Castillo-Víctor , M. E. ., Madrigal-Guridi , X., Martínez-Gómez , J. E. ., Torres-Hernández, E., Richter, L., Richter, S., Goolishian-Hernández , A. M. ., Mitchell, S., Haines, J. D., Dunlap-Smith, A. G., Place, E., Cox, C., Cox, R. ., Duarte-Ramos, F., & Sánchez-Ortiz , C. A. (2026). Los peces asociados a arrecifes del Parque Nacional Revillagigedo (Pacífico mexicano): un inventario curado, comentado e ilustrado. Revista Mexicana De Biodiversidad, 97, e975757. https://doi.org/10.22201/ib.20078706e.2026.97.5757